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! style="background: lightgreen;" | '''Plants'''
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Image:Adiantumpedatum.jpgFive-finger fern
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Scientific classification'''
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|Domain:||
Eukaryota
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|Kingdom:||'''Plantae'''
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!''''''Divisions''''''
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'''
Green algae'''
'''
land plants (embryophytes)'''
'''''non-vascular embryophytes'''''
Hepatophyta - liverworts
Anthocerophyta - hornworts
Bryophyta - mosses
'''
vascular plants (tracheophytes)'''
'''''seedless vascular plants'''''
Lycopodiophyta - clubmosses
Equisetophyta - horsetails
Pteridophyta – "true" ferns
Psilotophyta – whisk ferns
Ophioglossophyta - adderstongue ferns
'''''
seed plants (spermatophytes)'''''
Pinophyta - conifers
Cycadophyta - cycads
Ginkgophyta - ginkgo
Gnetophyta - gnetae
Magnoliophyta - flowering plants
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'''Plants''' are a major group of living things, including familiar organisms like trees, flowers, herbs, and ferns.
Aristotle divided all living things between plants, which generally do not move or have sensory organs, and animals. In
Linnaeus' system, these became the Kingdoms Vegetabilia (later Plantae) and Animalia. Since then, it has become clear that the Plantae as originally defined included several unrelated groups, and the
fungi and several groups of
algae were removed to new kingdoms. However, these are still often considered plants in many contexts. Indeed, any attempt to match "plant" with a single taxon is doomed to fail, because plant is a vaguely defined concept unrelated to the presumed phylogenic concepts on which modern taxonomy is based.
==Embryophytes==
:''See main article at
Embryophytes''
Most familiar are the
multicellular land plants, called
embryophytes. They include the
vascular plants, plants with full systems of
leaves,
stems, and
roots. They also include a few of their close relatives, often called ''bryophytes'', of which
mosses are the most common.
All of these plants have
eukaryotic cells with
cell walls composed of
cellulose, and obtain their energy through
photosynthesis, using light and
carbon dioxide to synthesize food. They are distinguished from
green algae, from which they evolved, by having specialized reproductive organs protected by non-reproductive tissues.
Bryophytes first appeared during the early
Palaeozoic. They can only survive in moist environments, and remain small throughout their life-cycle. This involves an alternation between two generations: a
haploid stage, called the
gametophyte, and a
diploid stage, called the
sporophyte. The sporophyte is short-lived and remains dependent on its parent.
Vascular plants first appeared during the
Silurian period, and by the
Devonian had diversified and spread into many different land environments. They have a number of adaptations that allowed them to overcome the limitations of the bryophytes. These include a cuticle resistant to desication, and vascular tissues which transport water throughout the organism. In many the sporophyte acts as a separate individual, while the gametophyte remains small.
Image:Spermatophyta.png Phylogeny of the modern Spermatophyta (seed plants) and some allied vascular plant groups. Note that the spore-bearing vascular plants are
paraphyletic with respect to the seed plants, with
ferns (Pteridophyta) more closely allied to seed plants than they are to
clubmosses (Lycopodiophyta)
The first primitive seed plants, Pteridosperms (seed ferns) and Cordaites, both groups now extinct, appeared in the late Devonian and diversified through the Carboniferous, with further evolution through the
Permian and
Triassic periods. In these the gametophyte stage is completely reduced, and the sporophyte begins life inside an enclosure called a
seed, which develops while on the parent plant, and with fertilisation by means of
pollen grains. Whereas other vascular plants, such as ferns, reproduce by means of spores and so need moisture to develop, some seed plants can survive and reproduce in extremely arid conditions.
Early seed plants are referred to as gymnosperms (naked seeds), as the seed embryo is not enclosed in a protective structure at pollination, with the pollen landing directly on the embryo. Four surviving groups remain widespread now, particularly the
conifers, which are dominant
trees in several
biomes. The angiosperms, comprising the
flowering plants, were the last major group of plants to appear, emerging from within the gymnosperms during the
Jurassic and diversifying rapidly during the
Cretaceous. These differ in that the seed embryo is enclosed, so the pollen has to grow a tube to penetrate the protective seed coat; they are the predominant group of flora in most biomes today.
==Algae and Fungi==
The
algae comprise several different groups of organisms that produce energy through photosynthesis. The most conspicuous are the
seaweeds, multicellular algae that often closely resemble terrestrial plants, found among the
green,
red, and
brown algae. These and other algal groups also include various single-celled creatures and forms that are simple collections of cells, without differentiated
tissues. Many can move about, and some have even lost their ability to photosynthesize; when first discovered, these were considered as both plants and animals.
The embryophytes developed from green algae; the two are collectively referred to as the green plants or Viridaeplantae. The kingdom Plantae is now usually taken to mean this
monophyletic group, as shown above. With a few exceptions among the green algae, all such forms have cell walls containing
cellulose and
chloroplasts containing
chlorophylls ''a'' and ''b'', and store food in the form of
starch. They undergo closed
mitosis without
centrioles, and typically have
mitochondria with flat cristae.
The chloroplasts of green plants are surrounded by two membranes, suggesting they originated directly from endosymbiotic
cyanobacteria. The same is true of the
red algae, and the two groups are generally believed to have a common origin. In contrast, most other algae have chloroplasts with three or four membranes. They are not in general close relatives of the green plants, acquiring chloroplasts separately from ingested or symbiotic green and red algae.
Unlike embryophytes and algae,
fungi are not photosynthetic, but are
saprophytes: they obtain their food by breaking down and absorbing surrounding materials. Most fungi are formed by microscopic tubes called hyphae, which may or may not be divided into cells but contain eukaryotic
nuclei. Fruiting bodies, of which
mushrooms are the most familiar, are actually only the reproductive structures of fungi. They are not related to any of the photosynthetic groups, but are close relatives of
animals.
==Importance==
The photosynthesis and carbon fixation conducted by land plants and algae are the ultimate source of energy and organic material in nearly all habitats. These processes also radically changed the composition of the Earth's
atmosphere, which as a result contains a large proportion of
oxygen. Animals and most other organisms are
aerobic, relying on oxygen; those that do not are confined to relatively few,
anaerobic environments.
Much of human nutrition depends on
cereals. Other plants that are eaten include
fruits,
vegetables,
herbs, and
spices. Some vascular plants, referred to as
trees and
shrubs, produce
woody stems and are an important source of building material. A number of plants are used decoratively, including a variety of
flowers.
==Growth==
Simple plants like algae may have short life spans as individuals, but their populations are commonly seasonal. Other plants may be organized according to their seasonal growth pattern:
*
Annual: live and reproduce within one growing season.
*
Biennial: live for two growing seasons; usually reproduce in second year.
*
Perennial: live for many growing seasons; continue to reproduce once mature.
Among the vascular plants, perennials include both
evergreens that keep their leaves the entire year, and
deciduous plants which lose their leaves for some part. In
temperate and
boreal climates, they generally lose their leaves during the winter; many
tropical plants lose their leaves during the dry season.
The growth rate of plants is extremely variable. Some mosses grow less than 1 μm/h, while most trees grow 25-250 μm/h. Some climbing species, such as
kudzu, which do not need to produce thick supportive tissue, may grow up to 12500 μm/h.
== Fossils ==
Plant
fossils include roots, wood, leaves, seeds, fruit,
pollen,
spores and
amber (the fossilized resin produced by some plants). Fossil land plants are recorded in terrestrial, lacustrine, fluvial and nearshore marine sediments.
Pollen,
spores and algae (
dinoflagellates and
acritarchs) are used for dating sedimentary rock sequences. The remains of fossil plants are not as common as fossil animals, although plant fossils are locally abundant in many regions worldwide.
Early fossil plants are well known from the
Devonian period, including the chert of Rhynie in Aberdeenshire,
Scotland. The best preserved examples, from which their cellular construction has been described, have been found at this locality. The preservation is so perfect that sections of these ancient plants show the individual cells within the plant tissue.
The
Coal Measures are a major source of
Palaeozoic plant fossils, with many groups of plants in existence at this time. The spoil heaps of coal mines are the best places to collect;
coal itself is the remains of fossilised plants, though structural detail of the plant fossils is rarely visible in coal. In the Fossil Forest at Victoria Park in
Glasgow,
Scotland, the stumps of ''Lepidodendron'' trees are found in their original growth positions.
The fossilized remains of conifer and angiosperm roots, stems and branches may be locally abundant in lake and inshore sedimentary rocks from the
Mesozoic and
Caenozoic eras.
Sequoia and its allies,
magnolia,
oak, and
palms are often found.
Petrified wood is common in some parts of the world, and is most frequently found in arid or desert areas were it is more readily exposed by erosion. Petrified wood is often heavily silicified (the organic material replaced by
silicon dioxide), and the impregnated tissue is often preserved in fine detail. Such specimens may be cut and polished using
lapidary equipment. Fossil forests of petrified wood have been found in all continents.
==References and further reading==
*Thomas N Taylor and Edith L Taylor. ''The Biology and Evolution of Fossil Plants''. Prentice Hall, 1993.
==See also==
*
Botany
*
Phytophilia (love of plants)
*
Vegetation Structure
==External links==
*
Tree of Life
===Botanical and Vegetation Databases===
*
United States of America
**
Eastern Oregon
*
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